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Submitted/ In Review/ In Revision

42. Reynolds, R.G., A.R. Puente-Rolón, J.P. Burgess, and B.O. Baker. A new boa from the Bahamas. submitted.

41. Reynolds, R.G., A.R. Puente-Rolón, A.L. Castle**, M. van de Schoot, and A. J. Geneva. Herpetofauna of the Cay Sal Bank, Bahamas and phylogenetic relationships of Anolis fairchildi, Anolis sagrei, and Tropidophis curtus from the region. submitted.

40. Quach*, Q.N., R.G. Reynolds, and L.J. Revell. Historical allopatry and secondary contact or primary intergradation in the Puerto Rican Crested Anole, Anolis cristatellus, on Vieques Island. in revision.


39. Reynolds, R.G., T.R. Strickland**, J.J. Kolbe, B.G. Falk, G. Perry, L.J. Revell, and J.B. Losos. Archipelagic genetics in a widespread Caribbean anole. Journal of Biogeography 44: 2631-2647. [doi] [Dryad]

38. Reynolds, R.G and S.T. Giery. Amphibians of the Bahamas and Turks and Caicos Islands. In: Heatwole, H., and N. Ríos-López. Amphibian Biology Vol. 9, Part 4. in press.

37. Reynolds, R.G. Invited book review: Natural History of Neotropical Treeboas (Genus Corallus). Copeia 105: 164-178. [doi]


36. Reynolds, R.G., S.T. Giery, W. Jesse*, and Q. Quach*. Preliminary assessment of road mortality in the Northern Bahamas Boa, Chilabothrus exsul. Caribbean Naturalist 34: 1-10. [doi] [pdf]

35. Reynolds, R.G., D.C. Collar, S.A. Pasachnik, M.L. Niemiller, A.R. Puente-Rolón, and L.J. Revell. Ecological specialization and morphological diversification in Greater Antillean boas.  Evolution 70: 1882-1895 [doi] [pdf] [Dryad]

34. Winchell*, K.M., R.G. Reynolds, S. Prado-Irwin**, A.R. Puente-Rolón, and L.J. Revell. Phenotypic shifts in urban areas in the tropical lizard Anolis cristatellus. Evolution 70: 1009-1022 [doi] [pdf] [Dryad] Cover Article

33. Reynolds, R.G., A.R. Puente-Rolón, K.J. Aviles-Rodriguez*,  A.J. Geneva, and N.C. Herrmann*. Discovery of a remarkable new boa from the Conception Island Bank, Bahamas. Breviora 549: 1-19 [doi]

32. Siliceo-Cantero, H.H., A. García, G. Pacheco, R.G. Reynolds, and B.C. Lister. Dimorphism and divergence in island and mainland anoles. Biological Journal of the Linnean Society 118: 852-872 [doi] [pdf]

31. Niemiller, M.L., B.M. Glorioso, D.B. Fenolio, R.G. Reynolds, S.J. Taylor, and B.T. Miller. Growth, survival, longevity, and population size of the Big Mouth Cave Salamander (Gyrinophilus palleucus necturoides) from the type locality in Grundy Co., Tennessee, USA. Copeia 2016: 35-41 [doi] [pdf]


30. Winchell*, K.M., P. Muralidhar*, J.R. Fredette**, and R.G. Reynolds. New herpetofaunal records for Great Isaac Cay, Bimini Group, Bahamas. IRCF Reptiles & Amphibians, Conservation and Natural History 22: 106-110. [doi] [pdf]

29. Reynolds, R.G., A.R. Puente-Rolón, R. Platenberg, R.K. Tyler**, P.J. Tolson, and L.J. Revell. 2015. Large divergence and low diversity suggest genetically informed conservation strategies for the endangered Virgin Islands boa (Chilabothrus monensis). Global Ecology and Conservation 3: 487-502 [doi] [pdf] [GitHub-R code]

28. Revell, L.J., D.L. Mahler, R.G. Reynolds, and G.J. Slater. 2015. Placing cryptic, recently extinct, or hypothesized taxa into an ultrametric phylogeny using continuous character data: a case study with the lizard Anolis roosevelti. Evolution 69: 1027-1035 [doi] [pdf]

27. Bushar, L.M., R.G. Reynolds, S. Tucker, L.C. Pace, W.I. Lutterschmidt, R.A. Odum, and H.K. Reinert. 2015. Genetic characterization of an invasive Boa constrictor population on the Caribbean island of Aruba. Journal of Herpetology 49: 602-610. [doi] [pdf] [Treebase]


26. Stuart, Y.E., T.S. Campbell, P.A. Hohenlohe, R.G. Reynolds, L.J. Revell, and J.B. Losos. 2014. Rapid evolution of a native species following invasion by a congener. Science 346: 463-466. [doi] [pdf] [Supplementary Materials]. Coverage in Washington Post, NY Times, The Scientist, Harvard Gazette, etc.

25. Pyron, R.A., R.G. Reynolds, and F.T. Burbrink. 2014. A taxonomic revision of boas (Serpentes: Boidae). Zootaxa 3846: 249-260 [doi] [pdf]

24. Reynolds, R.G., A.R. Puente-Rolón, M. Barandiaran, and L.J. Revell. 2014. Hispaniolan Boa (Chilabothrus striatus) on Vieques Island, Puerto Rico. Herpetology Notes 7: 121-122 [doi] [pdf]

23. Reynolds, R.G., M.L. Niemiller, and L.J. Revell. 2014. Toward a Tree-of-Life for the boas and pythons: multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201-213 [doi] [pdf][Treebase] [Curated Appendix B1]

22. Reynolds, R.G., A.R. Puente-Rolón, K. Kolodzaike**, and T. Butler-Smith**. 2014. Isolation and characterization of 23 novel polymorphic microsatellite markers from the endangered Puerto Rican boa (Chilabothrus inornatus) using paired-end Illumina shotgun sequencing. Conservation Genetics Resources 6: 107-109 [doi] [pdf]


21. Reynolds, R.G., M.L. Niemiller, S.B. Hedges, A. Dornburg, A.R. Puente-Rolón, and L.J. Revell. 2013. Molecular phylogeny and historical biogeography of West Indian boid snakes (Chilabothrus). Molecular Phylogenetics and Evolution 68: 461-470  [doi] [pdf] [Treebase] [Appendix A]

20. Puente-Rolón, A.R., R.G. Reynolds, and L.J. Revell. 2013. Preliminary genetic analysis supports cave populations as targets for conservation in the endemic endangered Puerto Rican boa (Epicrates inornatus). PLoS ONE   8(5): e63899  [doi] [pdf]

19. Reynolds, R.G. and B.M. Fitzpatrick. 2013.Tests of two methods for identifying founder effects in metapopulations reveal substantial type II error. Genetica 141: 119-131 [doi] [pdf] [GitHub- R code 1] [GitHub- R code 2]

18. Reynolds, R.G., A.R. Puente-Rolón, R.N. Reed, and L.J. Revell. 2013. Genetic analysis of a novel invasion of Puerto Rico by an exotic constricting snake. Biological Invasions 15: 953-959 [doi] [pdf] [Treebase]

17. Niemiller, M.L., J.R. McCandless, R.G. Reynolds, J. Caddle, T.J. Near, C.R. Tilquist, W.D. Pearson, and B.M. Fitzpatrick. 2013. Effects of climatic and geological processes during the Pleistocene on the evolutionary history of the northern cavefish, Amblyopsis spelaea (Teleostei: Amblyopsidae). Evolution 67: 1011-1025 [doi] [pdf]


16. Reynolds, R.G.,W. Booth, G.W. Schuett, B.M. Fitzpatrick, and G.M. Burghardt. 2012. Successive virgin births of viable male progeny in the checkered gartersnake, Thamnophis marcianus. Biological Journal of the Linnean Society 107: 566-572[doi] [pdf]

15. Reynolds, R.G., M.L. Niemiller, and B.M. Fitzpatrick. 2012. Genetic analysis of an endemic archipelagic lizard reveals sympatric cryptic lineages and taxonomic discordance. Conservation Genetics 13: 953-963[doi] [pdf]

14. Revell, L.J. and R.G. Reynolds. 2012. A new Bayesian method for fitting evolutionary models to comparative data with intraspecific variation. Evolution 66: 2697-2707[cover] [doi] [pdf]

13. Reynolds, R.G. and G.P. Gerber. 2012. Ecology and conservation of the Turks Island Boa (Epicrates c. chrysogaster: Squamata: Boidae) on Big Ambergris Cay. Journal of Herpetology  46: 578-586 [doi] [pdf]

12. Fitzpatrick, B.M., J.A. Fordyce, M.L. Niemiller, and R.G. Reynolds. 2012. What can DNA tell us about biological invasions? Biological Invasions 14: 245-253. [doi] [pdf]

11. Placyk, J.S., B.M. Fitzpatrick, G.S. Casper, R.L. Small, R.G. Reynolds, D.W.A. Noble, R.J. Brooks, and G.M. Burghardt. 2012. Hybridization between two gartersnake species (Thamnophis) of conservation concern: A threat or an important natural interaction? Conservation Genetics 13: 649-663. [doi] [pdf]

10. Buckner, S.D., R. Franz, and R.G. Reynolds. 2012. Bahama Islands and Turks & Caicos Islands. In: R. Powell and R.W. Henderson (Eds.). Island Lists of West Indian Amphibians and Reptiles. Bulletin of the Florida Museum of Natural History 51: 85-166. [doi] [pdf]

9. Reynolds, R.G. 2012. Epicrates chrysogaster. Catalogue of American Amphibians and Reptiles 898: 1-5. [pdf]


8. Reynolds, R.G., G.P. Gerber, and B.M. Fitzpatrick. 2011. Unexpected shallow genetic divergence in Turks Island Boas (Epicrates c. chrysogaster) reveals single evolutionarily significant unit for conservation. Herpetologica 67: 477-486. [doi] [pdf]

7. Booth, W., L. Million, R.G. Reynolds, G.M. Burghardt, E.L. Vargo, C. Schal, A.C. Tzika, and G.W. Schuett. 2011. Consecutive virgin births in the new world boid snake, the Colombian Rainbow Boa, Epicrates maurus. Journal of Heredity 102: 759-763. [doi] [pdf]

6. Reynolds, R.G. 2011. Status, conservation, and introduction of amphibians and reptiles in the Turks and Caicos Islands, British West Indies, p. 377-406. In: A. Hailey, B.S. Wilson, and J.A. Horrocks (Eds.). Conservation of Caribbean Island Herpetofaunas, vol. 2: Regional Accounts of the West Indies. Brill, Netherlands. [doi] [pdf]

5. Reynolds, R.G., M.L. Niemiller, and S.A. Pasachnik. 2011. Occurrence of pygmy salamanders (Desmognathus wrighti) in Monroe County, Tennessee. Journal of the Tennessee Academy of Science 86: 53-55. [pdf]

4. Niemiller, M.L., R.G. Reynolds, B.M. Glorioso, J. Spiess, and B.T. Miller. 2011. Herpetofauna of the cedar glades and adjacent habitats of the inner central basin of middle Tennessee. Herpetological Conservation and Biology 6: 135-149. [doi] [pdf]


3. Reynolds, R.G. and M.L. Niemiller. 2010. Island invaders: introduced amphibians and reptiles of the Turks and Caicos Islands. IRCF Reptiles & Amphibians: Conservation and Natural History 17: 117-121.


2. Miller, B.T., M.L. Niemiller, and R.G. Reynolds. 2008. Observations on egg-laying behavior and interactions among attending female Red Salamanders (Pseudotriton ruber) with comments on the use of caves by this species. Herpetological Conservation and Biology 3: 203- 210. [doi] [pdf]


1. Reynolds, R.G. and B.M. Fitzpatrick. 2007. Assortative mating in poison-dart frogs based on an ecologically important trait. Evolution 61: 2253-2259. [doi] [pdf]


Books (peer reviewed)


Niemiller, M.L., R.G. Reynolds, and B.T. Miller (Eds.). 2013. The Reptiles of Tennessee. The University of Tennessee Press, Knoxville. 366 pgs. [doi]




Niemiller, M.L. and R.G. Reynolds (Eds). 2011. The Amphibians of Tennessee. The University of Tennessee Press, Knoxville. 369 pgs. [doi]


Natural History Notes (peer reviewed):  <email me for reprints>

  1. Reynolds, RG, J. Burgess, G. Waters, and GP Gerber. Chilabothrus chrysogaster chrysogaster (Turks Island Boa). Diet. Herpetological Review accepted
  2. Reynolds, RG and AR Puente-Rolón. 2016. Chilabothrus strigilatus (Bahamas Boa). Distribution. Herpetological Review 47: 425.
  3. Reynolds, RG, NC Herrmann, AJ Geneva, AR Puente-Rolón, and KJ Aviles-Rodriguez. 2016. Osteopilus septentrionalis (Cuban Treefrog). Distribution. Herpetological Review 47: 420.
  4. Geneva, AJ, RG Reynolds, NC Herrmann, AR Puente-Rolón, and KJ Aviles-Rodriguez. 2016. Hemidactylus mabouia (Wood Slave). Distribution. Herpetological Review 47: 423-424.
  5. Reynolds, R.G. and B.N. Manco. 2013. Epicrates chrysogaster chrysogaster (Turks Island Boa). Climbing Behavior. Herpetological Review 44: 690-691.
  6. **Nelson, SK, ML Niemiller, and RG Reynolds. 2012. New county records for amphibians and reptiles in Tennessee. Herpetological Review 43: 353-354.
  7. Reynolds, RG. 2011. Hemidactylus mabouia (Tropical House Gecko), distribution. Caribbean Herpetology 28:1. [doi]
  8. Reynolds, RG. 2011. Eleutherodactylus planirostris planirostris (Cuban Flathead Frog), distribution. Caribbean Herpetology27:1. [doi]
  9. Reynolds, RG and BM Riggs. 2011. Iguana iguana (Green Iguana). Distribution. Herpetological Review 42: 241. [pdf]
  10. Reynolds, RG, ML Niemiller, BM Riggs, and BN Manco. 2011. Osteopilus septentrionalis (Cuban Treefrog). Distribution. Herpetological Review 42: 237-238. [pdf]
  11. Reynolds, RG and BM Riggs. 2011. Pantherophis guttatus (Corn Snake). Distribution. Herpetological Review 42: 243. [pdf]
  12. Reynolds, RG, BM Riggs, M Hibbert, and E Salamanca. 2011. Trachemys scripta elegans (Red-eared slider). Distribution. Herpetological Review 42: 239. [pdf]
  13. Reynolds, RG, J Batchasingh, and BM Riggs. 2011. Ramphotyphlops braminus (Brahminy Blind Snake). Distribution. Herpetological Review 42: 244. [pdf]
  14. Reynolds, RG, BM Riggs, M Hibbert, and E Salamanca. 2011. Anolis equestris (Knight Anole). Distribution. Herpetological Review 42: 239. [pdf]
  15. Reynolds, RG, ML Niemiller, and BN Manco. 2011. Epicrates chrysogaster chrysogaster (Turks Island Boa). Maximum Size. Herpetological Review 42: 290. [pdf]
  16. Reynolds, RG, and ML Niemiller. 2011. Epicrates chrysogaster chrysogaster (Turks Island Boa). Diet. Herpetological Review 42: 290. [pdf]
  17. Reynolds, RG, GP Gerber, and J Burgess. 2010. Tropidophis greenwayi greenwayi (Big Ambergris Dwarf Boa). Distribution. Herpetological Review 41: 520.
  18. Reynolds, RG. 2010. Typhlops platycephalus (Puerto Rican White-tailed Blindsnake), distribution. Caribbean Herpetology 15:1. [doi]
  19. Reynolds, RG, and ML Niemiller. 2010. Epicrates chrysogaster (Southern Bahamas Boa), distribution. Caribbean Herpetology 14:1. [doi]
  20. Reynolds, RG, and ML Niemiller. 2009. Hemidactylus mabouia (Wood Slave). Distribution. Herpetological Review 40: 452.
  21. Reynolds, RG. 2009. Leiocephalus psammodromus (Turks and Caicos Curly-tailed Lizard). Conspecific necrophagy. Herpetological Review 40: 88-89.
  22. Niemiller, ML, RG Reynolds, JG Reynolds, and BT Miller. 2009. Gyrinophilus porphyriticus porphyriticus (Northern Spring Salamander). Reproduction. Herpetological Review 40: 67.

Popular Articles:

  • Reynolds, RG and ML Niemiller. 2011. New Scales: Reptiles invade the Turks and Caicos. Green Pages, Times of the Islands Magazine, Spring, 94: 30-33. [pdf]
  • Reynolds, RG, and C Deal***. 2010. Where do all the babies go? Understanding the biology of juvenile Rainbow boas. Green Pages, Times of the Islands Magazine, Winter, 93: 34-36. [pdf]
  • Reynolds, RG. 2009. Scaling Millennia: What can lizards teach us about TCI’s history? Green Pages, Times of the Islands Magazine, Winter, 42-45. [pdf]
  • Reynolds, RG. 2008. The Snakes of the Turks and Caicos Islands. Brochure for the National Environmental Centre on Providenciales, Turks and Caicos Islands.

Blog Posts:

Technical Reports:

  • Miller, BT, D Estabrooks, RG Reynolds, ML Niemiller. 2009. Status review of the Streamside Salamanders (Ambystoma barbouri) in Tennessee. Report to Tennessee Wildlife Resources Agency. [pdf]
  • Reynolds, RG and ML Niemiller. 2009. Expedition report and recommendations for the Department of Environment and Coastal Resources. Unpublished Technical Report for the Ministry of Natural Resources, Turks and Caicos Islands.
  • Reynolds, RG. 2008. The reptiles of Ambergris Cay, British West Indies: report to staff naturalists. Unpublished Technical Report.

PDFs are for personal use only, and are made available especially for those without access to periodical resources. For all others, please use the [doi] links to go to the publisher’s webpage.